World Oldest San Diego Zoo Tortoise Gramma Dies at 141

141-Year-Old Galápagos Tortoise Gramma Passes Away at San Diego Zoo

Gramma, one of the oldest known Galápagos tortoises in captivity, has died at the age of 141 , marking the end of a remarkable life that spanned more than a century. A long-time resident of the San Diego Zoo , she was admired for her gentle temperament and became one of the institution’s most recognisable ambassadors for wildlife conservation.

A Century of Presence and Public Admiration

Originally from the Galápagos Islands, Gramma was later transferred to the United States and joined the San Diego Zoo after being housed previously at the Bronx Zoo. Historical accounts suggest her arrival took place around 1928–1931, making her one of the earliest animals in the zoo’s reptile collections. Over the decades, visitors remembered her slow but steady gait, calm nature, and towering shell figure — all of which contributed to her iconic status.

Final Years and Passing

In her later years, age-linked bone degeneration and mobility difficulties began affecting her quality of life. Following careful medical evaluation, zoo veterinarians made the compassionate decision to euthanise her on 20 November. Tributes poured in from former caretakers, zoo staff and long-time visitors, many of whom had seen her at various stages of their own lives.

Exam Oriented Facts

• Gramma lived to 141 years , among the longest on record for her species.

• She lived at San Diego Zoo for nearly a century.

• Galápagos tortoises can exceed 100–150 years , especially in captivity.

• 15 subspecies exist — several classified vulnerable to critically endangered .